MONA ISLAND

MARINE PROTECTED AREA

PUERTO RICO
 REEFKEEPER GOAL: To establish a marine protected area around Mona and Monito Islands, off southwest Puerto Rico.
 WHY: To protect the Island's pristine coral reefs from future development and from present anchor and overfishing damage.

Mona Marine Protected Area in Depth

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TO SUPPORT REEFKEEPER'S POSITION:

 CONTACT: Pedro A. Gelabert, Secretary
 AGENCY: Department of Natural and Environmental Resources
 ADDRESS: P.O. Box 5887
San Juan, PR 00906
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MONA AND MONITO ISLANDS
MARINE PROTECTED AREA
SITE NOMINATION
PUERTO RICO

ABSTRACT

Mona and Monito Islands are considered the Galápagos Islands of the Caribbean Sea. They are accessible only by boat. No other reef and offshore island habitat within U.S. jurisdiction possesses such invaluable habitat and biological diversity within such a reduced surface area. Their isolation from Puerto Rico and the Dominican Republic accounts for many unique biological interactions and endemic species within adjacent marine waters. Mona and Monito Islands have been recognized by the Commonwealth of Puerto Rico as a Natural Reserve for their ecological uniqueness, large biodiversity, including dozens of endemic species, and invaluable beauty. The site is also the critical habitat of endangered marine turtles, seabirds and occasional migratory marine mammals. Both islands are frequented by local commercial and recreational fishermen, divers and tourists. Mona supports extensive, well-developed coral reef spoor and groove systems, patch reefs, fringing reefs, drop-off communities, deep vertical walls, complex underwater caverns and rocky outcrops. Monito's bottom is characterized by vertical walls and deep rocky outcrops covered mainly by sponges and deepwater corals. Mona also has exuberant white sandy beaches and several shipwrecks which provide incentive for tourists and therefore make an important contribution to the regional economy. However, the coral reefs surrounding these islands are undergoing rapid degradation possibly due to overfishing, bioerosion, anchoring and ship grounding. In addition, there is recent increased interest in developing tourist facilities on Mona Island, which would significantly increase pressure on the pristine nature of the site's marine environment. There is at present no integrated management regime in place to protect the unique marine resources of the site. Therefore, ReefKeeper International is requesting the designation of Mona and Monito Islands as a Commonwealth Marine Protected Area for its protection and conservation.

Prepared by Edwin A. Hernandez-Delgado, M.S.

Nominated by ReefKeeper International Castillo del Mar -- Suite 1271 Isla Verde, Puerto Rico 00913 December 1, 1996

SITE LOCATION,

NAME AND DESCRIPTION

Site Name

Mona and Monito Islands, Puerto Rico

Location

Mona Passage, between Puerto Rico and the island of Santo Domingo. Latitude/Longitude: The area is located in a quadrangle within:

1. 18o10' N

2. 18o02'30"N

3. 67o57'30"W

4. 67o50' W

Mona and Monito are located within a faulting zone located at the Mona Passage halfway between the Commonwealth of Puerto Rico and the Dominican Republic. Mona is located over a 60.5 km2 (37.6 miles2) marine limestone platform which rises from depths of about 5,000 m (16,400 feet) on its southern face in the Caribbean Sea, and from about 458 m (1,500 feet) on the other sides. The two islands are separated by a narrow 220 m deep (720 feet) channel.

Mona and Monito Islands are accessible only by boat.

Description

Mona and Monito islands are 73.6 km (46 miles) off western Puerto Rico and 65 km (40 miles) off the eastern Dominican Republic (Cardona-Bonet, 1985a) (Figure 1). Mona's insular shelf, as defined by the 100-fathom (183 m or 600 feet) contour, covers only 5.6 km2 (3.5 miles2) or 9% of the total platform surface. Mona and Monito rise at the top of an underwater ridge which separates the Caribbean basin from the Atlantic Ocean. According to Cardona-Bonet (1985a), waters around the islands average between 366 and 1,159 m deep (1,200-3,800 feet). As a matter of fact, the deepest area of the Atlantic lies in waters just 142 km (88 miles) north of Mona, and is 8052 m deep (26,410 feet).

Extensive, well-developed coral reef spoor and groove systems, patch reefs, fringing reefs, drop-off communities, deep vertical walls, complex underwater caverns and rocky outcrops occupy most of the Mona Island insular shelf and attract many recreationists, divers and fishermen. These visitors are also attracted by Mona's white sandy beaches and shipwrecks. Monito's bottom is characterized by vertical walls and deep rocky outcrops covered mainly by sponges and deepwater corals.

Waters at the site are nearly pristine and crystal clear (Cintron et al., 1975), with horizontal visibility sometimes exceeding 46 to 61 m (150 to 200 feet). Moderate to strong currents come from the east, although there are some local fluctuations depending on bottom topography, tides, waves and wind (Perl and Cintron, 1974).

Mona Island's reef communities represent the most pristine, extensive and well-developed reefs in Puerto Rican waters. These are unusual in the variety of their formations because they include well-developed spoor and groove systems, drop-off communities, vertical walls, and extensive underwater caverns -- all of which supports an amazing biodiversity within a surface area of less than 5.6 km2 (3.5 miles2).

The site is also the critical habitat of endangered marine turtles, seabirds and occasional migratory marine mammals. No other reef and offshore island habitat within U.S. jurisdiction possess such invaluable habitat and biological diversity within such a reduced surface area.

CURRENT STATUS

Mona and Monito Islands are a Natural Reserve under the Puerto Rico Department of Natural Resources (DNR). However, management measures only address upland natural resources. The existing regulatory regime does not adequately protect Mona marine resources from the increasing pressure of human activities.

RATIONALE FOR CONSIDERATION

AS A COMMONWEALTH MARINE PROTECTED AREA

A. The site's marine biodiversity is both unique to and the largest of the region.

B. Mona and Monito isolation from Puerto Rico and the Dominican Republic accounts for many unique biological interactions and endemic species. They are considered the Galápagos Islands of the Caribbean Sea.

C. The reefs are undergoing rapid degradation possibly due to overfishing, bioerosion, anchoring and ship grounding.

D. Mona and Monito Islands are a discrete site conducive to management.

E. Mona supports extensive and well-developed coral formations surrounded by pristine marine waters.

F. Both islands possess extensive and well-developed drop-off communities and vertical walls with many deepwater species occurring within safe diving limits.

G. Several endangered marine turtles make use of the sandy beaches, reefs and surrounding areas. Migratory endangered marine mammals can be frequently seen in Mona's surrounding waters.

H. The reefs provide incentive for tourist visits by boat and therefore make an important contribution to the regional economy.

I. There is recent increased interest in developing tourist facilities on Mona Island, which would significantly increase pressure on the pristine nature of the site's marine environment.

J. Mona and Monito Islands have been recognized by the Commonwealth of Puerto Rico as a Natural Reserve.

K. Mona has been the focus of many research activities.

Site Uniqueness and Particular Ecosystem Values

Mona Island marine resources are characterized by a diverse array of tropical marine ecosystems, including: patch reefs, fringing reefs, spur and groove systems, underwater caverns, drop-off communities, vertical walls, abundant black corals and deep water sponges, hard bottoms, rocky outcrops, algal reefs, sandy beaches, shipwrecks and a highly rich biodiversity.

Biodiversity and Species Richness and Uniqueness

Mona Island supports an amazing biodiversity which includes: at least 23 scleractinian coral species (Cintrón et. al., 1975), dozens of marine invertebrates (E. Ortiz, 1973; Hernández-Delgado, 1993), dozens of marine algae (A. Ortiz, 1973), over 270 species of marine fishes (Prentice, 1973), 417 plant species -- 8 endemics of Mona and 27 of this region of the Caribbean, plus 78 rare or endangered in Puerto Rico (Woodbury et al., 1974), 10 types of vegetational associations (Rogers and Cintrón, 1974), about 20 endangered animals (DNR, 1975), 16 terrestrial mollusks -- including 4 endemics (Velez, 1970), nearly 100 marine mollusks (Mestey, 1991), 51 spiders (Velez, 1971), at least 526 insects -- including 24 endemics (Martorell, 1973), 13 reptiles and 1 endemic amphibian (Rivero, 1978; Wiewandt, 1973), nearly 97 bird species -- with 2 endemics on Mona (Biaggi, 1983), and 9 terrestrial mammals -- 4 of which are native (Wiewandt, 1973).

No other single reef habitat and offshore island within U.S. jurisdiction possesses such invaluable habitat and biological diversity within such a reduced surface area.

Biological Productivity

The large, well-developed and complex coral reefs and other benthic communities are the outstanding marine biological feature of this area. Mona is inhabited by scleractinian corals, octocorals, precious corals, hydrozoans, sponges, spiny lobsters, octopuses, queen conches, echinoderms, worms, shells, endangered marine turtles, mammals, birds, and fishes. These latter include: sharks, rays, barracudas, groupers, squirrelfishes, snappers, grunts, jacks, goatfishes, angelfishes, damselfishes, wrasses, trunkfishes, filefishes, butterflyfishes, parrotfishes, doctorfishes, dolphins, mackerels, tuna, flyingfishes and others. This is one of the most productive habitats in the region.

Species Maintenance

The many habitats of this site provide different biological uses for many different species. This includes nesting and feeding areas for endangered marine turtles, as well as feeding areas for marine birds. These habitats also provide areas for feeding, reproduction, shelter and juvenile development and growth for many other species, including those of commercial consumptive value.

Habitat Features and Uniqueness

Mona Island and Monito possess an exuberant and unique diversity of underwater habitats which are briefly described below:

1. Spur and groove systems

These are characterized by a complex array of grooves through the forereef and spurs rising from the sandy bottom which are oriented parallel to prevalent strong currents and surge (Cintron et al., 1975; Canals et al., 1981). Major spur and groove development occur off Mona in the south, between Playa Pájaros and Punta Caigo o No Caigo, and in the west between Cabo Barrio Nuevo and Punta Oeste (Figure 2). These areas possess the most extensive reef development, and are characterized by the highest underwater biological diversity, and highest coral and sponge cover. Coral cover ranges from 6 to 80% (Canals et al., 1981). These areas also support the largest diversity of reef fishes. Their aesthetic and recreational value is unmeasurable. The near absence of an insular shelf at Monito prevents major reef development.

2. Fringing and patch reefs

Extensive development of shallow fringing, patch and rock reefs occurs on Mona from Uvero, in the south coast, to Sardinera in the west. These reefs are characterized by relatively sparse coral growth, brown algae, sponges and a variety of other epibenthic fauna. Coral cover rarely exceeds 30% at Uvero and 10% at Sardinera (Canals et al., 1981), although Hernández-Delgado (1993) reported an average coral cover of 29% at a small patch reef just north of Sardinera. This area is a very important nursery area for the growth, feeding and shelter of many juvenile reef fishes and invertebrates. Turbidity is slightly higher due to strong surge. This area is not adequate for most recreational purposes because of prevalent conditions and inaccessibility from both camping areas and from water. However, the sponges within this habitat area are very important food resources for the endangered green turtle Eretmochelys imbricata, which is almost exclusively spongivorous. The shorelines of these areas are also used as nesting sites by E. imbricata and the leatherback turtle Dermochelys coriacea (Hernandez-Guzman, in press).

3. Carabinero drop-off

The Carabinero drop-off is located in the southern coast of Mona Island (Figure 2) and rivals the drop-offs of La Parguera (southwestern Puerto Rico) and other locations in the Caribbean. It is characterized by a 180 m (590 feet) wide and less than 10 m deep (33 feet) insular platform. The insular shelf drops off at about this depth. At 15 m (49 feet) there is a 5 m (16 feet) wide underwater terrace which represents an ancient sea level terrace. Similar terraces can be observed at 38 and 40 m (124 and 131 feet). Then the shelf drops off to 76 m (250 feet) where there is another narrow terrace. It drops off further to depths of 800 m (more than 2,700 feet). The Carabinero drop-off boasts about 23 scleractinian coral species, an anthipatarian (black coral) forest and large sponge diversity (Cintrón et al., 1975). According to Canals et al. ( 1981), live coral cover at 10 m (33 feet) averages 18%, and only 7.4% at 20 m (66 feet) at the drop-off. There, macroalgae and sponges become abundant, with 21% and 24% cover, respectively. Octocorals, reef fishes and marine invertebrates are also abundant. This is perhaps the most frequented fishing ground off Mona, particularly by hook-and-liners and ocassional spearfishers. It is aesthetically invaluable for experienced divers. It is believed that Monito is also characterized by nearly similar drop off communities as Mona although there is no published data.

4. Las Carmelitas underwater cavern system

Las Carmelitas is a complex submerged cavern system located at the north of Playa Sardinera, western Mona Island, lying within the western spoor and groove reef system (Figure 3). Its origin seems to be the result of the collapse of large limestone blocks from ancient cliffs or reef rocks (Cintrón et al., 1975). It averages depths shallower than 6 m (20 feet). Then it drops seaward to about 18 m (60 feet). Coral cover ranges from 12% to 37%. The giant basket sponge Xestospongia muta becomes abundant near the drop-off at 30 m (98 feet). This is one of the most amazing and unique Puerto Rican underwater resources. It also provdies shelter for a myriad of reef fishes and cryptic creatures. However, marine currents can become occasionally dangerous and strong, particularly during winter months. This is the second most important fishing ground and diving spot off Mona.

5. Deep vertical walls

Mona possesses 30.8 km (19.1 miles) of shoreline, but 17.6 km (10.9 miles) of them, or 57.2%, are vertical cliffs. These are located in the south, east, north and northwest coasts of the island. Most of these cliffs plunge vertically to an average of 30 m (98 feet) down to the sandy bottom (Hendrick, 1973). The northern vertical walls are characterized also by the presence of collapsed enormous limestone blocks which support well-developed brown algal forests and sparse scleractinian coral, octocoral, black coral, hydroid and sponge growth (Cintrón et al., 1975). These habitats are commonly frequented by the endangered sea turtle E. imbricata (Kontos, 1986). Sharks can be commonly seen from the top of the cliffs. Large commercially important fishes (e.g. snappers, groupers, jacks) are also common at these sites (Hendrick, 1973). Water transparency can reach more than 46-61 m (150-200 feet). These deep vertical walls have not been recreationally exploited because strong, treacherous open water currents, bottom depth and lack of safe anchoring areas discourage recreational diving and fishing. However, hook-and-line fishing is common. These ecosystems are unique within Puerto Rican and U.S. waters. Monito Island is completely surrounded by vertical cliffs which plunge vertically into deep waters. They must also support deep wall communities similar to those at Mona.

6. Algal reefs

Mona Island also supports outstanding and well developed algal-dominated reefs located just south-southwest of Punta Arenas, in the southwest coast. The rocky bottom is frequently smothered by sediments moved by strong and unpredictable currents, which prevents major coral growth. The bottom is covered mainly by brown algae (e.g. Sargassum spp., Turbinaria spp.). These are very important grounds for juveniles organisms and grazers. Recreational use is non-existent due to the unpredictable currents and strong surge regime.

7. Rocky shores and hardbottoms.

Rocky shores, outcrops and hardbottoms are among the most common type of bottom habitats at Mona. They support sparse populations of corals, sponges, algae and many epibenthic invertebrates. These habitats are also very important grounds for the development and growth of juvenile individuals of many important species.

8. Pelagic waters.

Pelagic waters surrounding Mona and Monito support frequent heavy international ship traffic, which is responsible for the presence of occasional oil tars. Nevertheless, these waters are frequented by endangered marine turtles (Erdman, 1973). In addition, migratory marine mammals can be commonly seen in the Mona Passage. According to Philibosian and Yntema (1977), at least five marine mammal species frequent these waters. These are the big whale Zyphiuscavirostris, whale Physeter catodon, humpback whale Megapteranovaeangliae, dolphin Globicephala macrorhyncha and saddleback dolphin Delphinus delphis. Mona's pelagic waters are also home to several commercially important pelagic fishes such as dolphins, tuna, mackerels, marlins, sharks, etc.

Historical and Cultural Resources

Mona possesses valuable underwater archeological resources. According to Cerame Vivas (1976), there are about 11 known shipwrecks around the island. These include some Hispanic galleons. However, most shipwrecks have been poorly documented (Cardona-Bonet, 1985b).

In past centuries, Mona Island was the home of "Taino" indians and earlier cultures. Later, it was used as a base for smuggling and pirate attacks in the Caribean region. During the last century and until recent decades, Mona supported "Guano" mining activities and a small resident population. There are many historical sites in Mona's coastal plains and cave systems (e.g. archeological sites and objects, petroglyphs). Mona was also the target of U.S. Navy military activities and bombing, as well as target of a German submarine attack during the Second World War.

HUMAN USES AND IMPACTS

Fishery Resources and Overfishing

Mona and Monito Islands support a significant portion of the western Puerto Rico artisanal fisheries, particularly for snappers, groupers and lobsters. Most of the fishery comes from hook-and-lining of both reef and pelagic fishes. However, fishtrapping is commonly employed around south-southwestern Monito. Extensive spearfishing is common at shallow and moderately deep reefs around Mona.

Mona and Monito are heavily overfished by both Puerto Rican and Dominican fishermen. The latter is an illegal practice, since the Dominicans are fishing within Federal and Puerto Rican State waters. Endangered marine turtles are also commonly fished illegally around Mona and Monito. However, at present there is no on-site law enforcement personnel to enforce existing regulations.

A recent evaluation of marine resources at Sardinera, western Mona, made by Hernandez-Delgado (1993), identified 63 reef fish species. The most common and abundant groups were herbivore fishes such as surgeonfishes (Acanthuridae) and black durgons Melichthys niger. Parrotfishes (Scaridae) were also common. However, there were basically no significant predator species. No groupers (Serranidae) and adult snappers (Lutjanidae) were seen although these groups were fairly common at this locality about 10 years ago (Hernandez-Delgado, personal observations). These highly prized predator groups are now restricted to deeper waters near the insular shelf drop-off. This may be the result of overfishing.

According to Hernandez-Delgado (1993), densities of the corallivorous molluscan gastropod Coralliophila abbreviata appear to have increased at Sardinera. The densities and feeding activity of this snail upon Acropora spp. (elkhorn and staghorn) corals could increase as a result of decreased mollusc predation in the reef areas (Brawley and Adey, 1982). This situation may be the indirect result of snapper-grouper overfishing, since it coincides with a low number of predators in reef areas. In addition, most of the dead coral heads and the bases of many live corals (e.g. Acroporapalmata, Diploria spp. and Siderastreasiderea) are being overgrown by the boring sponge Cliona spp. and by the demosponge Chondrilla nucula. According to Hernandez-Delgado (1993), sponge cover in a small fringing reef at Sardinera ranged between 20 and 90%, with an average cover of 63%. Crustose and filamentous algal cover ranged from 5 and 90%, averaging 36%.

Recreational Resources and Associated Impacts

Mona and Monito have high aesthetic value and are highly prized for recreational diving. They are accessible only by boat. Snorkeling, SCUBA diving, underwater photography, fishing (including spearfishing), and occasional pleasure boating are common in the area.

There are no mooring buoy systems at Mona or Monito. All types of vessels anchor directly over reef or seagrass bottoms, causing in many instances irreversible damage to bottom communities.

During the 1980's, two large vessels -- the "Alborada" (1980) and the "A Regina" (1985) -- ran aground at Playa Pájaros on Mona. There is no significant information about the ecological consequences of the Alborada grounding. However, waters around the shipwreck -- which is stii there -- are usually murky and live coral cover has decreased almost to 0%. The ecological impacts of the "A Regina" accident have been well documented by Bohnsack and Harper (1985), and by Cintrón et. al. (1987). These impacts included a shift in reef fish community composition, reductions in water transparency and coral cover, direct destruction of corals, and coral smothering by sedimentation.

Military Use Impacts

The area was formerly used as a military ordnance testing ground. There are no military uses at present.


Mineral Resources and Ocean Mining

From the 1800's to mid 1900's, Mona Island supported "guano" mining activities. Several historical sites remain almost intact on the island. No mineral extraction has occurred in decades.

Island Development and Associated Marine Impacts

There has been recent pressure to develop Mona for tourism, which may be a threat to the pristine nature of the site. There is already an unpaved airport on Mona, although at present it is closed to the public. There are plans to establish an expanded, paved runway. Other possible projects may include the construction of small tourist villas, a dock, a road, and possibly sewage treatment facilities. The Commonwealth of Puerto Rico also has plans to sell electricity in the near future to the Dominican Republic, and the main underwater powerline is supposed to pass through Mona Island.

PURPOSE AND NEED FOR DESIGNATION

While the marine habitats off Mona and Monito Islands have so far withstood man-induced pressures, such success can not realistically be expected in the future without deliberate protection. The existing regulatory regime does not adequately protect the site's unique marine resources from the increasing pressure of human activities.

A marine protected area management plan for Mona and Monito Islands could address its protection from excessive human impacts through:

1) regulations to prevent damage to resources and habitats, e.g. coral damage due to anchoring, drilling, mining, fishing gear, and live rock collection; archeo- logical resource loss due to unregulated salvaging; and irreparable loss of biodiversity and community stability due to extreme overfishing,

2) provision of the long-term planning and management necessary to protect the site's irreplaceable habitats and ecosystems, and

3) establishment of a resource assessment program to monitor the health of the site's marine communities, research the site's uniquely isolated biological communities, and provide information needed for future management decisions.

There is no comprehensive program for the long-term protection, assessment and management of the unique Mona and Monito Islands marine ecosystem. Designation as a Commonwealth Marine Protected Area would provide the means to correct such deficiencies. The unique characteristics of these islands' marine habitats should of themselves warrant their designation as a Marine Protected Area. However, when one adds to this uniqueness the increased man-induced pressures possible from fishing, mining, drilling, anchoring and collection, the designation of these islands as Commonwealth Marine Protected Areas becomes imperative.

# # #

Literature Cited

Biaggi, V. 1983. Las aves de Puerto Rico, Edicion Revisada y Aumentada. EditorialUniversitaria, Universidad de Puerto Rico, Rio Piedras, P.R. 378 pp.

Bohnsack, J.a., & D.E. Harper. 1985. Reef Fish survey, Mona Island, Puerto Rico, October, 1985. Report Submitted to the Puerto Rico DDepartment of Natural Resources, San Juan, P.R. 16pp.

Brawley, S.H., and W.H. Adey. 1982. Coralliophila abbreviata: A significant corallivore. Bull. Mar. Sce. 32(2):595-599.

Canals, M., H. Ferrer, y H. Merced. 1981. Los arrecifes de coral de Isla de Mona. 1- 26. Octavo Smposio de Recursos Naturales, 7 de diciembre de 1981. Departamento de Recursos Naturales, San Juan, P.R.

Cardona-Bonet, W.A. 1985a. Islotes de Borinquen: (Amona, Abey, Pinas, Sikeo y otros). Notas para su historia. Oficina Estatal de Preservacion Historica de La Fortaleza, San Juan, P.R.

Cardona-Bonet, W.A. 1985b. Los naufragios de Puerto Rico. Arqueologia (Referencia desconecida):16-18.

Cerame-Vivas, M.J. 1976. Marine Atlas of Puerto Rico. (Xerox copy):2-30.

Cintron, G., J. Thurston, J. Williams, y F. Mackenzie. 1975. Caracteristicas de la plataforma insular de Isla de Mona. 69-91. Segundo Simposio de Recursos Naturales, 26 de agosto de 1975. Departamento de Recursos Naturales, San Juan, P.R.

Cintron-Molero, G., R. Martinez, & B.B. Cintron. 1987. Effects of the "A Regina" grounding on coral reefs of Mona Island, Puerto Rico, Final Report. Coastal Zone Management Program Task No. 7.1, Department of Natural Resources, San Juan, P.R. 72pp + App.

Department of Natural Resources. 1975. Unit Plan for the Management of the Mona Island Forest Reserve. Forestry Task Force, Department of Natural Resources, San Juan, P.R.

Diez, C.E., R. van Dam, & V.P. Vicente. In press. Estudio preliminar de la ecologia alimentaria del carey de concha (Eretmochelys imbricata) en Isla de Mona, Puerto Rico. XIX Simposio de los Recursos Naturales, 14 de Octubre de 1993, Departament de Recursos Naturales, San Juan, P.R.

Erdman, D.S. 1973. Pelagic life around Mona Island. E-1 - E-3. In F. Pagan-Font (Ed.), Las islas de Mona y Monito. Una evaluacion de sus Recursos Naturales e Historicos, Vol. 2, Junta de Calidad Ambiental, San Juan, P.R.

Hendrick, W. 1973. Underwater World of Mona. F-3 - F-6. In F. Pagan-Font (Ed.), Las islas de Mona y Monito. Una evaluacion de sus Recursos Naturales e Historicos, Vol. 2, Junta de Calidad Ambiental, San Juan, P.R.

Hernandez-Delgado, E.A. 1993. Informe preliminar sobre el estado de los recursos marinos de Isla de Mona y algunas recomendaciones de manejo. Informe preliminar sometido a la Comision de Recursos Naturales y Calidad Ambiental, Camara de Representantes, Estado Libre Asociado de Puerto Rico, San Juan, PR, 18 de marzo de 1993. 42pp + App.

Hernandez-Guzman, G.M. In press. Tortugas marinas en la Reserva Natural de Isla de Mona, Puerto Rico, 1992. XIX Simposio de los Recursos Naturales, 14 de Octubre de 1993, Departament de Recursos Naturales, San Juan, P.R.

Kontos, A.R. 1986. Sea turtle research report, 1985 Mona Island, Puerto Rico. U.S. Fish and Wildlife Service, Ecological Services. 1-40 + tables + maps + graphs.

Martorell, L.F. 1973. The insects of Mona Island, Puerto Rico. J-1 - J-7. In F. Pagan-Font (Ed.), Las islas de Mona y Monito. Una evaluacion de sus Recursos Naturales e Historicos, Vol. 2, Junta de Calidad Ambiental, San Juan, P.R.

Mestey, V. 1991. Los moluscos mas comunes de Isla de Mona. Naturaleza. Feb. 1991: 36-38

Ortiz, A. 1973. Marine flora. F-7 - F-8. In F. Pagan- Font (Ed.), Las islas de Mona y Monito. Una evaluacion de sus Recursos Naturales e Historicos, Vol. 2, Junta de Calidad Ambiental, San Juan, P.R.

Ortiz, A. 1973. Marine Invertebrates. F-8 - F-9. In F. Pagan-Font (Ed.), Las islas de Mona y Monito. Una evaluacion de sus Recursos Naturales e Historicos, Vol. 2, Junta de Calidad Ambiental, San Juan, P.R.

Perl, H.W., & G. Cintron. 1974. Preliminary findings: Nearshore surface currents in the vicinity of Sardinera and Uvero, Mona Island: December-January 1973-74. 73-80. Primer Simposio de Recursos Naturales, 29 de agosto de 1974. Departamento de Recursos Naturales, San Juan, P.R.

Philibosian, R., & J.A. Ynterna. 1977. Annotated checklist of the birds, mammals, reptiles and amphibians of the Virgin Islands & Puerto Rico: 1-48

Prentice J. 1973. Fish Resource. F-9 - F-10. In F. Pagan-Font (Ed.), Las islas de Mona y Monito. Una evaluacion de sus Recursos Naturales e Historicos, Vol. 2, Junta de Calidad Ambiental, San Juan, P.R.

Raffaelle, H. 1973. Assessment of Mona Island avifauna. K-1 - K-32. In F. Pagan-Font (Ed.), Las islas de Mona y Monito. Una evaluacion de sus Recursos Naturales e Historicos, Vol. 2, Junta de Calidad Ambiental, San Juan, P.R.

Rivero, J.A. 1978. Los anfibios y reptiles de Puerto Rico. Editorial Universitaria, Universidad de Puerto Rico, Rio Piedras, P.R. 152pp.

Rogers, L. & B. Cintron. 1974. Plant communities of Mona Island. Pittman Robinson Project W-817 Study III, Job-1, Final Report. 1-18.

Velez, M.J. 1970. Preliminary checklist of the terrestrial and freshwater fauna of the greater Puerto Rico region. Mimeograph. Dept. of Biology, University of Puerto Rico, Rio Piedras, P.R. 1-71.

Velez, M.J. 1971. Key to the spiders fauna of Puerto Rico. Mimeograph. Dept. of Biology, University of Puerto Rico, Rio Piedras, P.R. 1-39.

Vicente,V.P. 1993. Habitos alimentarios del carey de concha (Eretmochelys imbricata). 150-156. XVIII SImposio de Recursos Naturales, 17 y 18 de noviembre de 1992, Departamento de Recursos Naturales, San Juan, P.R.

Wiewandt, T. A. 1973. Mona amphibians, reptiles and mammals. L-1- L-13. In F. Pagan-Font (Ed.), Las islas de Mona y Monito. Una evaluacion de sus Recursos Naturales e Historicos, Vol. 2, Junta de Calidad Ambiental, San Juan, P.R.

Woodbury, R., L. Martorell, & J. Garcia-Tuduri. 1977. The flora of Mona and Monito Island, Puerto Rico (West Indies). Estacion Experimental Agrícola, Bulletin 252, Rio Piedras, P.R. 1-9.

Mona Marine Protected Area in Depth

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